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This U.S.S. Enterprise-A shuttle was commandeered by Sybok on Nimbus III. With Kirk and his landing party hostage, and with Sulu at the helm, Sybok forced Kirk to take him to Enterprise, so that the zealous Vulcan could use the starship to reach Sha Ka Ree.

Kirk instructed Sulu and the Enterprise to prepare for "Emergency Landing Plan 'B'," which eventually lead to crashing the shuttle in the Enterprise's hangar deck.

Uhura also used the shuttle to pick up Kirk and other crew members from Yosemite National Park.